COURSE SYLLABUS

SYMBIOSIS

1	Course Title:	SYMBIOSIS
2	Course Code:	BIO5105
3	Type of Course:	Optional
4	Level of Course:	Second Cycle
5	Year of Study:	1
6	Semester:	1
7	ECTS Credits Allocated:	6
8	Theoretical (hour/week):	3
9	Practice (hour/week) :	0
10	Laboratory (hour/week) :	0
11	Prerequisites:	There is no prerequisite for this course.
12	Recommended optional programme components:	None
13	Language:	Turkish
14	Mode of Delivery:	Face to face
15	Course Coordinator:	Prof. Dr. ŞABAN GÜVENÇ
16	Course Lecturers:	Prof.Dr. Şule ÖZTÜRK
17	Contactinformation of the Course Coordinator:	Prof.Dr. Şaban GÜVENÇ E-mail:saban@uludag.edu.tr Telefon: +90 (224) 2941793 Adres: Uludağ Üniversitesi, Fen – Edebiyat Fakültesi, Biyoloji Bölümü, Görükle Kampüsü, 16059 Nilüfer/Bursa.
18	Website:
19	Objective of the Course:	Aim of this course is to understand and discriminate the major symbiosis types. The morphological and anatomical structures of the two most common symbiosis examples (mycorrhizae and lichens) and the importance of physiological and ecological aspects of their is understand.
20	Contribution of the Course to Professional Development	The morphological and anatomical structures of Mycorrhiza and Lichens, the two most common common life types, contribute to their professional development by understanding their physiological and ecological importance.

Learning Outcomes:

1	To understand the types and examples of symbiosis in nature.;
2	To know the nature of lichens and to understand the importance of lichens in ecosystem;
3	To knowledge and skills for the separation of lichen symbionts and to use appropriate techniques.;
4	To acquire the skill of reading, understanding and evaluating articles in related fields with the help of terminology and information learned from the study.;
5	To gain the skill to apply their thesis studies to the knowledge, experience and discipline gained from this course.;
6	To be able to adapt information acquired on life in marine and other aquatic environments to other disciplines and to work and take part in cooperative projects.;

22	Course Content:

Week	Theoretical	Practical
1	Types of symbiotic relationship (mutualism, parasitism and commensalism).
2	Endosymbiont: Mycorrhiza
3	Morphological and anatomical features of lichen thallus.
4	Lichen classification.
5	Reproductive structures of lichenized fungi.
6	The symbiotic phenotype, evolution of cyanolichens fossil lichens, possible relations with mycorrhizal symbioses
7	Cyanolichens: diversity of fungal-cyanobacterial associations, mycobionts and photobionts of cyanolichens, structural - functional organisation of cyanolichens, types of cyanolichens
8	Repeating courses and midterm exam
9	Carbon metabolism in cyanolichens: carbon metabolism and the poikilohydric life-style, soluble carbohydrates and secondary metabolites, respiration and photosynthesis
10	Nitrogen metabolism in cyanolichens: heterocysts, glutamine synthetase, nitrogen fixation, transfer of fıxed nitrogen and partitioning among the partners, pathways of nitrogen metabolism
11	Origin and evolution of green lichen algae: phylogeny within the genus trebouxia, the most common lichen alga, lichenized trebouxia.
12	Lichenized trentepohlia
13	The importance of the functional aspects of lichen symbiosis
14	The methods used to separate lichen partners.

23	Textbooks, References and/or Other Materials:	Seckbach, J. Symbiosis: Mechanisms and Model Systems. Kluwer Academic Publishers, 2004. Ahmadjian, V. The Lichen Symbiosis. John Wiley & Sons. Inc., New York, 1993. Thomas H. Nash III. Lichen Biology. Cambridge University Press., Cambridge, 1996.
24	Assesment

TERM LEARNING ACTIVITIES	NUMBER	PERCENT
Midterm Exam	1	40
Quiz	0	0
Homeworks, Performances	0	0
Final Exam	1	60
Total	2	100
Contribution of Term (Year) Learning Activities to Success Grade		40
Contribution of Final Exam to Success Grade		60
Total		100
Measurement and Evaluation Techniques Used in the Course	The system of relative evaluation is applied
Information	It is evaluated to what extent students reach the course gains / outcomes with written exams in measurement and evaluation.

ECTS / WORK LOAD TABLE

CONTRIBUTION OF LEARNING OUTCOMES TO PROGRAMME QUALIFICATIONS

LO: Learning Objectives

PQ: Program Qualifications

Contribution Level:

1 Very Low

2 Low

3 Medium

4 High

5 Very High